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Werner Schwarzhans

    A comparative morphological treatise of recent and fossil otoliths of the family Sciaenidae (Perciformes)
    A comparative morphological treatise of recent and fossil otoliths of the order Pleuronectiformes
    The otoliths from the miocene of the North Sea basin
    Otolithen aus den Gerhartsreiter Schichten (Oberkreide: Maastricht) des Gerhartsreiter Grabens (Oberbayern)
    Fish otoliths from the Paleocene of Bavaria (Kressenberg) and Austria (Kroisbach and Oiching-Graben)
    Otoliths from dredges in the Gulf of Guinea and off the Azores
    • 2012

      Otoliths from 54 teleost species are reported from Paleocene strata in Bavaria and Austria, covering 31 families, with 33 new species described and 7 in open nomenclature. Until recently, descriptions of Paleocene otolith assemblages were scarce, but knowledge has expanded to over 100 valid species, making this collection the largest and most diverse. Kressenberg mainly represents a neritic shelf fauna, while Kroisbach includes open marine elements, particularly Stomiiformes, marking the first such association from the Paleocene. The regional diversification of Paleocene otolith-based fish faunas is notable, with data from the U.S.A., Belgium, Denmark, Greenland, and Ukraine enabling a paleobiogeographic evaluation. This collection is significant for its transitional nature in Teleostei evolution, showcasing forms emerging post-K-T boundary extinction and before modern Teleostei arose in the Eocene after the PETM event. Few Paleocene species overlap with Late Cretaceous or Eocene species. The Paleocene assemblage features a low level of extinct morphologies and an abundance of plesiomorphic ones, including modern groups like Percoidei, Congridae, Ophidiiformes, and Stomiiformes. Gadiformes are underrepresented, while Beryciformes, once dominant in the Late Cretaceous, remain common in the Paleocene of Bavaria. Many taxa are considered 'extinct plesiomorphic' or 'missing links' in teleost phylogeny. Phylogenetic analyses and in

      Fish otoliths from the Paleocene of Bavaria (Kressenberg) and Austria (Kroisbach and Oiching-Graben)
    • 2010

      Inhaltsverzeichnis1. Einleitung 7 2. Oberkreide-Otolithen (eine Bestandsaufnahme) 9 3. Systematischer Teil 14 3.1 Artenliste 15 3.2 Beschreibung der Arten 16 3.2.1 Ordnung Osteoglossiformes 16 3.2.2 Ordnung Elopiformes 16 3.2.3 Ordnung Anguilliformes 21 3.2.4 Ordnung Siluriformes 26 3.2.5 Ordnung Salmoniformes 26 3.2.6 Ordnung Stomiiformes 30 3.2.7 Ordnung Aulopiformes 33 3.2.8 Ordnung Myctophiformes 36 3.2.9 Ordnung Ophidiiformes 38 3.2.10 Ordnung Beryciformes 46 3.2.11 Ordnung Zeiformes 68 3.2.12 Ordnung Perciformes 73 3.2.13 Ordnung Scorpaeniformes 81 3.2.14 Otolithen incertae sedis und Lapilli 81 4. Faunistische Rekonstruktion 84 4.1 Entwicklung der Teleosteer anhand von Otolithen in Mittel-Europa 84 4.2 Faunenzusammensetzung und Vergleich mit Skelettfunden 85 4.3 Palökologie 86 5. English Summary and Phylogenetic Conclusions 87 5.1 Phylogenetic usage of otoliths 87 5.2 Methodology and limitations of otolith usage 87 5.3 Functional morphology 87 5.4 General aspects of the Maastrichtian otolith fauna from Bavaria 89 5.5 Faunal reconstruction 90 5.6 Phylogenetic interpretations 90 6. Literatur 99

      Otolithen aus den Gerhartsreiter Schichten (Oberkreide: Maastricht) des Gerhartsreiter Grabens (Oberbayern)
    • 2010

      This monograph describes and reviews otoliths from the Miocene of the North Sea Basin, based on around 90,000 specimens spanning nearly the entire Miocene stratigraphic range. Over half of these specimens are from the Reinbekian and Hemmoorian stages, equivalent to the Early Serravallian to Late Burdigalian. It recognizes 269 species and five subspecies, including 32 extant species recorded as fossils, 65 new species, three new subspecies, and 32 species in open nomenclature. A primary goal is to document otoliths through photographs and consistent definitions, aiming to serve as a reference handbook for fossil otoliths from the region. The analysis reveals that the Early Miocene Vierlandian fauna closely resembles the Late Oligocene Chattian fauna. The Hemmoorian and Reinbekian stages show an abundance of new species adapted to warmer climates, with some also found in the Aquitaine Basin and Paratethys. Species diversity is high, with pairs suggesting allopatric speciation due to marine conduit openings. The Hemmoorian and Reinbekian stages likely connected the North Sea Basin to the subtropical NE-Atlantic and possibly the northern Paratethys. A significant faunal shift occurs in the Langenfeldian, marked by the immigration of temperate species. The analysis indicates a lifespan of 3 to 15 million years for most species, with some lasting up to 32 million years. The otolith database allows for a critical review of the biostr

      The otoliths from the miocene of the North Sea basin