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The narrative follows Cambridge professor Horace Holly and his ward Leo Vincey as they embark on an adventure to a lost African kingdom, spurred by a mysterious package revealing secrets about Leo's ancestry. Their journey leads them to encounter a primitive tribe and the enigmatic white queen Ayesha, who embodies both power and a deep connection to Leo. The story delves into late Victorian themes of race, evolution, and female authority, sparking both admiration and critique for its portrayal of womanhood.